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Here two individuals have always to be combined, and the pedigree becomes a more complicated one. Such is the case with the toad-flax, which is nearly sterile with its own pollen. But even in these cases the visits of insects bringing pollen from other plants, must be carefully excluded.
In the spring, a starling comes to his nest in a cleft of the cliff above; he shoots over from the dizzy edge, spreads his wings, borne up by the ascending air, and in an instant is landed in his cave. On the sward above, in the autumn, the yellow lip of the toad-flax, spotted with orange, peers from the grass as you rest and gaze how far? out upon the glorious plain.
Even the red valerian, that sprouts so boldly in bushes of coral from the top of the wall, is regarded by some people as a weed and an impudent intruder. For myself, I love the spectacle of stone walls breaking out into flower with red valerian and ivy-leaved toad-flax. The country people have greeted these flowers with comic and friendly names.
They could never be overlooked, on the other hand, in experimental culture. The first record of the peloric toad-flax is that of Zioberg, a student of Linnaeus, who found it in the neighborhood of Upsala. This curious discovery was described by Rudberg in his dissertation in the year 1744.
The step from the ordinary toad-flax to the peloric form is short, and it appears as if it might be produced by slow conversion. The ordinary species produces from time to time stray peloric flowers. These occur at the base of the raceme, or rarely in the midst of it. In other species they are often seen at the summit. Terminal pelories are usually regular, having five equal spurs.
Such facts clearly point to a common origin, and as only the terminal flowers are affected by the malformation, the fertility of the whole plant is evidently not seriously infringed upon. Before leaving the labiates, we may cite a curious instance of pelorism in the toad-flax, which is quite different from the ordinary peloric variety.
The ground for this choice lies simply in the fact that the peloric toad-flax is known to have originated from the ordinary type at different times and in different countries, under more or less divergent conditions. It had arisen from time to time, and hence I presumed that there was a chance to see it arise again.
Broad discs of greater knapweed with stalks like wire, and yellow toad-flax with spotted lip grow among it. Grasping this tough grass as a handle to climb up by, the explorer finds a rising slope of sward, and having walked over the first ridge, shutting off the road behind him, is at once out of civilisation. There is no noise.
For they named a lovely little lilac snapdragon, Linaria Domini Pellicerii "Lord Pellicier's toad-flax;" and that name it will keep, we may believe, as long as winter and summer shall endure. But to return.
The peloric toad-flax in my experiment was seen to arise thrice from the same strain. Three different individuals of my original race showed a tendency to produce peloric mutations, and they did so in a number of their seeds, exactly as the mutations of the evening-primroses were repeated nearly every year.
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