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Updated: May 24, 2025
All these cases are intelligible on the hypothesis of pangenesis; for they depend on the gemmules of certain parts, although present in both sexes, becoming, through the influence of domestication, either dormant or developed in either sex.
Palmaris accessorius, muscle variations of the. Pampas, horses of the. Pangenesis, hypothesis of. Panniculus carnosus. Pansch, on the brain of a foetal Cebus apella. Papilio, proportion of the sexes in North American species of; sexual differences of colouring in species of; coloration of the wings in species of. Papilio ascanius. Papilio Sesostris and Childrenae, variability of. Papilio Turnus.
I read the chapter on Pangenesis first, for I could not wait. I can hardly tell you how much I admire it. It is a positive comfort to me to have any feasible explanation of a difficulty that has always been haunting me, and I shall never be able to give it up till a better one supplies its place, and that I think hardly possible.
Yours, as far as I can see, explains everything in growth and reproduction, though of course the mystery of life and consciousness remains as great as ever. Parts of the chapter on Pangenesis I found hard reading, and have not quite mastered yet, and there are also throughout the discussions in Vol.
According to the theory of Pangenesis, it is necessary to assume that all the gemmules of this anomalous formation, and therefore the mother-gemmule of the cell, and the daughter-gemmules of the special epidermic tissue, and of the very singular subjacent tissue of the endothecium, have been perpetuated, and transmitted from parent to offspring in a dormant state, and through a number of generations, such as startles the imagination, and leads it to refuse its consent to the theory of Pangenesis, however seductive it may be."
A somewhat paradoxical outcome of the speculations of those who profess to rely exclusively on the testimony of sense. "Les extrêmes se touchent," and extreme sensationalism shakes hands with the "das seyn ist das nichts" of Hegel. Altogether the hypothesis of Pangenesis seems to be little, if at all, superior to anterior hypotheses of a more or less similar nature.
Yet on the hypothesis of Pangenesis, the zoospore of an alga must contain gemmules from all the cells of the parent algæ, and from all the parts of all their less remote ancestors in all their stages of existence. What wonder then that such an excessively complex body should divide and multiply; and what parity is there between such a body and a gemmule?
In the second volume the causes and laws of variation, inheritance, etc., are discussed as far as our present state of knowledge permits. Towards the end of the work I give my well-abused hypothesis of Pangenesis.
Thus, then, there are grave difficulties in the way of the reception of the hypothesis of Pangenesis, which moreover, if established, would leave the evolution of individual organisms, when thoroughly analysed, little if at all less mysterious or really explicable than it is at present.
Still we cannot exclude the possibility of such a transmission occasionally occurring, for even if the greater part of the effects must be attributable to natural selection, there might be a smaller part in certain cases which depends on this exceptional factor." I repeatedly tried to understand Mr. Darwin's theory of pangenesis, and so often failed that I long since gave the matter up in despair.
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