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Updated: May 24, 2025


Apart from the atoms of Democritus, and apart also from the speculations of mediæval writers, the molecules of Bonnet and of Buffon almost anticipated the hypothesis of Pangenesis.

"Thus, in Pangenesis, everything proceeds by force of unknown elements, and we may ask whether it is more logical to prefer a system which assumes a multitude of unknown elements to a system which assumes only a single one?" Mr. Darwin appears, by "Natural Selection," to destroy the reality of species, and by Pangenesis that of the individual. Mr.

In addition to the theory of "Natural Selection," by which it has been attempted to account for the origin of species, Mr. Now, though the hypothesis of Pangenesis is no necessary part of "Natural Selection," still any treatise on specific origination would be incomplete if it did not take into consideration this last speculation of Mr. Darwin.

"But, it may be urged in opposition by the Pangenesists, your vital principle is an unknown and irresolute x. This is true; but, on the other hand, let us see whether Pangenesis produces a clearer formula, and one free from unknown elements.

Weismann's theory is clear, simple and convenient, but incomplete; for, unlike Darwin's theory of pangenesis, it scarcely attempts any real explanation of the extremely complex potentialities possessed by the reproductive elements.

Towards the end of the work is propounded the hypothesis of Pangenesis, which has been generally rejected, and which the author himself looked upon as unverified, although by it a remarkable number of isolated facts could be connected together and rendered intelligible. The "Descent of Man" was published in February, 1871.

I have nothing, however, to do with Mr. Darwin's theory of pangenesis beyond avoiding the pretence that I understand either the theory itself or what Professor Weismann says about it; all I am concerned with is Professor Weismann's admission, made immediately afterwards, that the somatic cells may, and perhaps sometimes do, impart characteristics to the germ-cells.

Among the additions I intended making while the book was in the press, was a chapter on Mr. Darwin's provisional theory of Pangenesis, which I felt convinced must be right if it was Mr. Darwin's, and which I was sure, if I could once understand it, must have an important bearing on "Life and Habit." I had not as yet seen that the principle I was contending for was Darwinian, not Neo- Darwinian.

I am sure all naturalists will be disgusted at the malicious and ignorant article in the Athenæum. It is a disgrace to the paper, and I hope someone will publicly express the general opinion of it. We can expect no good reviews of your book till the quarterlies or best monthlies come out.... I shall be anxious to see how Pangenesis is received. Believe me yours very faithfully,

Still we cannot exclude the possibility of such a transmission occasionally occurring, for even if the greater part of the effects must be attributable to natural selection, there might be a smaller part in certain cases which depends on this exceptional factor." I repeatedly tried to understand Mr. Darwin's theory of pangenesis, and so often failed that I long since gave the matter up in despair.

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