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Of certain characteristics of song nothing more; and therefore to suppose that it furnishes a complete explanation, which satisfies all the requirements of scientific logic, of so wonderful an intonation as that, for example, of the Marsh-Warbler, or that no other relationships, except that of the territory, enter into the total emotional complex, simplifying here or elaborating there to meet the exigencies of diverse circumstances to suppose this would be foolish.
The small plantation or wooded bank may hold a Nightingale one year, but we miss its song there the next; the osier bed or gorse-covered common which vibrates with the trill of the Grasshopper-Warbler one April is deserted the following season; the plantation which is occupied by a host of common migrants this summer may be enlivened next year by the song of the rarer Marsh-Warbler also; and so on.
For example, the Grasshopper-Warbler, Marsh-Warbler, Nightingale, Corncrake, Red-backed Shrike, or Whinchat have each some distinctive peculiarity which makes them conspicuous, and each one is subject to marked fluctuation in numbers.
What could be more unlike than the songs of the Willow-Warbler and of the Chiffchaff, of the Marsh-Warbler and the Reed-Warbler, or of the Yellow Bunting and the Cirl Bunting?
The Marsh-Warbler can utter the call of the Green Woodpecker, or sing as the Nightingale does, with as much facility as it sings its own song; and the Blackcap is well-nigh as proficient in copying the cries and melodies of surrounding species and so, if it were necessary, we might proceed to add to the list.
The plaintive notes of the Willow-Warbler and of the Chiffchaff are to our ears very closely akin, so, too, are those of the Marsh-Warbler and of the Reed-Warbler, and there is a great resemblance between the hissing sound produced by the two Whitethroats. In Co.
But just why a Marsh-Warbler is gifted with a voice that is so beautiful and varied, whilst the Grasshopper-Warbler must perforce remain content with a monotonous trill; just why the tail feathers of the Snipe have developed into an instrument, whilst the Pied Woodpecker has developed muscles which enable it to make use of a decayed branch as an instrument we know no more than we do of the nature of the forces which lead the Reed-Warbler to weave its nest to reeds, or the caterpillar of the Elephant Hawk Moth to assume so peculiar an attitude when disturbed.
Districts frequented by the Marsh-Warbler and offering plenty of situations of the type required by the bird are often inhabited by a few members only, and yet the disposition to remain in a definite position is just as marked.
More striking still is the change in the case of the Marsh-Warbler, and the sudden deterioration, or even suspension, of strains so beautiful and so varied, at a moment, too, when it might least be expected, at once arrests the attention.
This, it seems to me, is the purpose of the peculiar call of the female a call which, so far as biological interpretation is concerned, is just as much a song as the melody of the Marsh-Warbler and its interest for us just now lies in this, that here we have a special case in which the sexes have separate territories, the female is polyandrous, and the voice of the male is not sufficient by itself to bring to pass the union of the sexes; and in which, consequently, if the purpose of song be that of recognition, we should expect to find, as we do find, that the female had a distinct and penetrating call.
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