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Updated: May 4, 2025


I believe it has been unconsciously used; and thus only can I understand the several rules and guides which have been followed by our best systematists. As we have no written pedigrees, we are forced to trace community of descent by resemblances of any kind.

This fact was, of course, most distasteful to the systematists of his time and even for a long period afterwards they attempted to discredit it. Milde and many others have opposed these new ideas with some temporary success. Only of late has the school of Jordan received due recognition, after Thuret, de Bary, Rosen and others tested its practices and openly pronounced for them.

Hence modifications of structure, viewed by systematists as of high value, may be wholly due to the laws of variation and correlation, without being, as far as we can judge, of the slightest service to the species.

Such very rich genera however, are not the rule, but are exceptional cases, indicating the rarity of powerful mutative changes. On the other hand, species may remain in a state of constancy during long, apparently during indefinite, ages. Many facts plead in favor of the constancy of species. This principle has always been recognized by systematists.

I believe it has thus been unconsciously used; and only thus can I understand the several rules and guides which have been followed by our best systematists. We have no written pedigrees; we have to make out community of descent by resemblances of any kind.

The same holds good in many other cases, and systematists have often become uncertain as to the true value of some form, by its relationship to the allied types in the way of retrogressive modification. Color-differences are so showy, that they easily overshadow other characters.

Wallace, or when analogous views on the origin of species are generally admitted, we can dimly foresee that there will be a considerable revolution in natural history. Systematists will be able to pursue their labours as at present; but they will not be incessantly haunted by the shadowy doubt whether this or that form be a true species.

It should be remembered that systematists are far from pleased at finding variability in important characters, and that there are not many men who will laboriously examine internal and important organs, and compare them in many specimens of the same species.

Intermediate forms occur, connecting one group with another in a manner sadly perplexing to systematists, except to those who have ceased to expect absolute limitations in Nature. All this blending could hardly fail to suggest a former material connection among allied forms, such as that which an hypothesis of derivation demands.

I believe it has thus been unconsciously used; and only thus can I understand the several rules and guides which have been followed by our best systematists. We have no written pedigrees; we have to make out community of descent by resemblances of any kind.

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