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Updated: May 11, 2025


And the knowledge we now possess justifies us completely in the anticipation, that when the still lower Eocene deposits, and those which belong to the cretaceous epoch, have yielded up their remains of ancestral equine animals, we shall find, first, a form with four complete toes and a rudiment of the innermost or first digit in front, with probably a rudiment of the fifth digit in the hind foot; while, in still older forms, the series of the digits will be more and more complete, until we come to the five-toed animals, in which, if the doctrine of evolution is well founded, the whole series must have taken its orgin.

For example, as we study the history of the horse family we find that an originally five-toed animal began to walk more and more on its middle toe, in such a way that this toe received more and more use, while the outer toes were used less and less.

The proboscidians culminated in the Pliocene, when some of the giant elephants reached a height of fourteen feet. THE ARTIODACTYLS comprise the hoofed Mammalia which have an even number of toes, such as cattle, sheep, and swine. Like the perissodactyls, they are descended from the primitive five-toed plantigrade mammals of the lowest Eocene.

We may generally distinguish three groups among the Vertebrates in relation to the formation of their limbs. The third group comprises the four higher classes of Vertebrates the amphibia, reptiles, birds, and mammals; in these quadrupeds there are at first the same two pairs of limbs, but in the shape of five-toed feet.

Among the older incidents in the evolution of organic life, the changes were very wonderful which out of the pectoral fin of a fish developed the jointed fore-limb of the mammal with its five-toed paw, and thence through much slighter variation brought forth the human arm with its delicate and crafty hand.

In the cloven-hoofed forms, a single toe first disappeared, leaving but four; then the two outer of these were aborted, leaving two nearly equal digits. In the series of the horse, where we can trace the change more clearly, we find the earliest form five-toed, but the outer and inner digit shrunken so as to become of little use.

And the knowledge we now possess justifies us completely in the anticipation, that when the still lower Eocene deposits, and those which belong to the Cretaceous epoch, have yielded up their remains of ancestral equine animals, we shall find, first, a form with four complete toes and a rudiment of the innermost or first digit in front, with, probably, a rudiment of the fifth digit in the hind foot; while, in still older forms, the series of the digits will be more and more complete, until we come to the five-toed animals, in which, if the doctrine of evolution is well founded, the whole series must have taken its origin.

If the expectation raised by the splints of the Horses that, in some ancestor of the Horses, these splints would be found to be complete digits, has been verified, we are furnished with very strong reasons for looking for a no less complete verification of the expectation that the three-toed. Plagiolophus-like "avus" of the horse must have had a five-toed "atavus" at some earlier period.

We now believe that polled cattle, five-toed Dorkings, top-knotted Houdans, frizzles and black skinned chickens arose through mutations. Burbank's Methods The wonderful Burbank with his thornless cactus, his stoneless plum, and his white blackberry, is simply a searcher after mutations. His success is not because he uses any secret methods, but because of the size of his operations.

We find in the Carboniferous period fossilised remains of five-toed amphibia, the oldest terrestrial, air-breathing vertebrates. These amphibia increase in variety in the Permian epoch. Towards the close of it we find the first Amniotes, the ancestors of the three higher classes of Vertebrates.

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