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The Brachiopods have passed into entirely new and more advanced species in the many advances and retreats of the shores, but the Molluscs show more interesting progress. The commanding group from the start is that of the Molluscs which have "kept their head," the Cephalopods, and their large shells show a most instructive evolution.
He set his screws in motion, and kept them circling in the vortices thus created until a boat could be lowered. And these specimens were whole cephalopods and fragments of cephalopods, some of gigantic proportions, and almost all of them unknown to science!
The highest group, the Cephalopods, "keep their heads," in the literal sense, and we shall find them advancing from form to form until, in the octopus of a later age, they discard the ancestral shell, and become the aristocrats of the Mollusc kingdom. The last and most important line that led upward from the chaos of Archaean worms is that of the Arthropods.
The whole series has been divided by zones characterised by particular Ammonites; for while other families of shells pass from one division to another in numbers varying from about 20 to 50 per cent, these cephalopods are almost always limited to single zones, as Quenstedt and Oppel have shown for Germany, and Dr. Wright and others for England.
The pretty nautilus is the only survivor to-day of the vast Mesozoic population of coiled-shell Cephalopods. A rival to the Ammonite appeared in the Triassic seas, a formidable forerunner of the cuttle-fish type of Cephalopod. The animal now boldly discards the protecting and confining shell, or spreads over the outside of it, and becomes a "shell-fish" with the shell inside.
A curious resemblance, though less in degree, has also been seen to exist between the auditory organs of fishes and of Cephalopods. Remarkable similarities between certain placental and implacental mammals, between the bird's-head processes of Polyzoa and the pedicellariæ of Echinoderms, between Ichthyosauria and Cetacea, with very many other similar coincidences, have also been pointed out.
Yet if we compare the older Reptiles and Batrachians, the older Fish, the older Cephalopods, and the eocene Mammals, with the recent members of the same classes, we must admit that there is truth in the remark. Let us see how far these several facts and inferences accord with the theory of descent with modification.
In the complex struggle for life it is quite credible that crustaceans, not very high in their own class, might beat cephalopods, the highest molluscs; and such crustaceans, though not highly developed, would stand very high in the scale of invertebrate animals, if judged by the most decisive of all trials the law of battle.
Among Cephalopods, the Family of Squids contains several Genera distinguished by the structure of the solid shield within the skin of the back, by the form and connection of their fins, by the structure of the suckers with which their arms are provided, by the form of their beak, etc.
But the highest class of Mollusks, the Cephalopods or Chambered Shells, or Cuttle-Fishes, as they are called when the animal is unprotected by a shell, are, on the contrary, very well preserved, and they are very numerous. Of these I will speak somewhat more in detail, because their geological history is a very curious one.
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