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Updated: July 1, 2025
To put it metaphorically, the mammary gland and the uterus, after the change of life, the lip, after the decay of the teeth, have done their work, outlived their usefulness, and are being placed upon a starvation pension by a grateful country.
Unless indeed we confine the term "primary" to the reproductive glands, it is scarcely possible to decide which ought to be called primary and which secondary. The female often differs from the male in having organs for the nourishment or protection of her young, such as the mammary glands of mammals, and the abdominal sacks of the marsupials.
In 1894 Richard Semon further proved that these large eggs, rich in food-yelk, have a partial segmentation and discoid gastrulation, as I had hypothetically assumed in 1879; here again they resemble their reptilian ancestors. The construction of the mammary gland is also peculiar in the Monotremes.
Similarly with the mammary glands. It is not an unreasonable supposition that by the effects of greater or less function, inherited through successive generations, these may be enlarged or diminished in size; but it is out of the question to allege such a cause for changes in their numbers.
A similar, but still stronger, argument may be based upon the existence of teats, and even functional mammary glands, in male mammals. Numerous cases of "Gynaecomasty," or functionally active breasts in men, are on record, though there is no mammalian species whatever in which the male normally suckles the young.
It is regarded of especial moment that the mirror, taken as a whole, be symmetrical, and especially that the mammary mirror be so; yet it often occurs that it is far otherwise, its outline being often very fantastical exhibiting deep bays, so to speak, and islands of downward growing hair.
The mammary glands of the Ornithorhynchus may, perhaps, be considered, in comparison with the udder of a cow, as in a nascent state. The ovigerous frena of certain cirripedes, which are only slightly developed and which have ceased to give attachment to the ova, are nascent branchiæ.
He unbuttoned successively in reversed direction waistcoat, trousers, shirt and vest along the medial line of irregular incrispated black hairs extending in triangular convergence from the pelvic basin over the circumference of the abdomen and umbilicular fossicle along the medial line of nodes to the intersection of the sixth pectoral vertebrae, thence produced both ways at right angles and terminating in circles described about two equidistant points, right and left, on the summits of the mammary prominences.
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