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Updated: May 25, 2025


It cannot be maintained that species when intercrossed are invariably sterile, and varieties invariably fertile; or that sterility is a special endowment and sign of creation.

These two kinds of flowers sometimes differ wonderfully in structure, yet may be seen to graduate into each other on the same plant. The ordinary and open flowers can be intercrossed; and the benefits which certainly are derived from this process are thus secured.

Right and left it went, this silent gallery, and although he was unaware of the fact, it joined other like galleries which encircled the slopes and met and intercrossed so that one might wander for hours along these mystic aisles of the hills. Below again, beyond a sloping woody thicket, lay the meadows and farmlands sweeping smoothly onward to the heath.

He hid his canoe in the bushes and strode rapidly across the islet, pushing with impatience through the twigs of heavy undergrowth intercrossed over his path. From motives of prudence he would not take his canoe to the meeting-place, as Nina had done. He had left it in the main stream till his return from the other side of the island.

The view generally entertained by naturalists is that species, when intercrossed, have been specially endowed with the quality of sterility, in order to prevent the confusion of all organic forms. This view certainly seems at first probable, for species within the same country could hardly have kept distinct had they been capable of crossing freely.

The fertility of varieties, that is of the forms known or believed to have descended from common parents, when intercrossed, and likewise the fertility of their mongrel offspring, is, on my theory, of equal importance with the sterility of species; for it seems to make a broad and clear distinction between varieties and species.

It cannot be maintained that species when intercrossed are invariably sterile and varieties invariably fertile; or that sterility is a special endowment and sign of creation.

The view generally entertained by naturalists is that species, when intercrossed, have been specially endowed with the quality of sterility, in order to prevent the confusion of all organic forms. This view certainly seems at first probable, for species within the same country could hardly have kept distinct had they been capable of crossing freely.

I have as yet spoken as if the varieties of the same species were invariably fertile when intercrossed. But it is impossible to resist the evidence of the existence of a certain amount of sterility in the few following cases, which I will briefly abstract. The evidence is at least as good as that from which we believe in the sterility of a multitude of species.

Nevertheless the perfect fertility of so many domestic varieties, differing widely from each other in appearance, for instance of the pigeon or of the cabbage, is a remarkable fact; more especially when we reflect how many species there are, which, though resembling each other most closely, are utterly sterile when intercrossed.

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