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Updated: July 3, 2025
And in the widespread Arctogaeal province, the Pliocene and later mammals belong to the same groups as those which now exist in the province. The law of succession of types, therefore, holds good for the present epoch as compared with its predecessor. Does it equally well apply to the Pliocene fauna when we compare it with that of the Miocene epoch?
But if we now cross the frontier between the Cainozoic and the Mesozoic faunae, as they are preserved within the Arctogaeal area, we meet with an astounding change, and what appears to be a complete and unmistakable break in the line of biological continuity.
But if there was this Australian facies about both the terrestrial and the marine faunae of Mesozoic Europe, and if there is this unaccountable and immense break between the fauna of Mesozoic and that of Tertiary Europe, is it not a very obvious suggestion that, in the Mesozoic epoch, the Australian province included Europe, and that the Arctogaeal province was contained within other limits?
In fact, as I have recently endeavoured to prove to the Society, there was an Arctogaeal continent and an Arctogaeal province of distribution in Triassic times as there is now; and the Sauropsida and Marsupialia which constituted that fauna were, I doubt not, the progenitors of the Sauropsida and Marsupialia of the whole Mesozoic epoch.
Looking at the present terrestrial fauna of Australia, it appears to me to be very probable that it is essentially a remnant of the fauna of the Triassic, or even of an earlier, age ; in which case Australia must at that time have been in continuity with the Arctogaeal continent. But now comes the further inquiry.
But it is very remarkable that while the Miocene fauna of the Arctogaeal province, as a whole, is of the same character as the existing fauna of the same province, as a whole, the component elements of the fauna were differently associated.
But it is very remarkable that while the Miocene fauna of the Arctogaeal province, as a whole, is of the same character as the existing fauna of the same province, as a whole, the component elements of the fauna were differently associated.
By great good fortune, an extensive mammalian fauna of the latter epoch has now become known, in four very distant portions of the Arctogaeal province which do not differ greatly in latitude.
But Austro-Columbia presents difficulties from which Australia is free; Cantelidoe and Tapirdoe are now indigenous in South America as they are in Arctogaea; and, among the Pliocene Austro-Columbian mammals, the Arctogaeal genera Equus, Mastodon, and Machairodus are numbered. Are these Postmiocene immigrants, or Praemiocene natives? Cunningham sent over to me some time ago from Patagonia.
The Permian epoch marks the commencement of a new movement of upheaval in our area, which attained its maximum in the Triassic epoch, when dry land existed in North America, Europe, Asia, and Africa, as it does now. Into this great new continental area the Mammals, Birds, and Reptiles developed during the Palaeozoic epoch spread, and formed the great Triassic Arctogaeal province.
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