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So again, it is probable, from what we know of the embryos of mammals, birds, fishes and reptiles, that these animals are the modified descendants of some ancient progenitor, which was furnished in its adult state with branchiae, a swim-bladder, four fin-like limbs, and a long tail, all fitted for an aquatic life.

Again, an organ may become rudimentary for its proper purpose, and be used for a distinct object: in certain fish the swim-bladder seems to be rudimentary for its proper function of giving buoyancy, but has become converted into a nascent breathing organ or lung. Other similar instances could be given.

This assures the zoölogist that the origin of the lungs in the higher animals is found in the swim-bladder of the so-called lungfish. In this Silurian time certain of these lungfish were perhaps trapped in the basin in the marsh by the uplifting of the border.

Again, an organ may become rudimentary for its proper purpose, and be used for a distinct object: in certain fish the swim-bladder seems to be nearly rudimentary for its proper function of giving buoyancy, but has become converted into a nascent breathing organ or lung. Other similar instances could be given.

All physiologists admit that the swim-bladder is homologous, or "ideally similar" in position and structure with the lungs of the higher vertebrate animals: hence there is no reason to doubt that the swim-bladder has actually been converted into lungs, or an organ used exclusively for respiration.

The oxygen in the air was slowly killing him, and yet his swan song was possible because of an inner atmosphere so rich in this gas that it would be unbreathable by a creature of the land. Nerve and muscle, special expanse of circling bones, swim-bladder and its tenuous gas all these combined to produce the aquatic harmony.

At a still earlier period the progenitors of man must have been aquatic in their habits; for morphology plainly tells us that our lungs consist of a modified swim-bladder, which once served as a float. The clefts on the neck in the embryo of man shew where the branchiae once existed.

This subject is intimately connected with that of the gradation of the characters, often accompanied by a change of function, for instance, the conversion of a swim-bladder into lungs, points which were discussed in the last chapter under two headings. Nevertheless, I will here consider in some detail several of the cases advanced by Mr.

All physiologists admit that the swim-bladder is homologous or "ideally similar" in position and structure with the lungs of the higher vertebrate animals; hence there seems to me to be no great difficulty in believing that natural selection has actually converted a swim-bladder into a lung, or organ used exclusively for respiration. p. 191.

In the cases in which we know of no intermediate or transitional states, we should be extremely cautious in concluding that none can have existed, for the metamorphoses of many organs show what wonderful changes in function are at least possible. For instance, a swim-bladder has apparently been converted into an air-breathing lung.