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Updated: May 4, 2025
Contrasted with these, Artiodactyla have always an even number of functional digits, the third and fourth reaching the ground symmetrically, bearing the weight and forming the "split hoof;" the second and fifth remain, in most cases, as mere vestiges, showing externally as the accessory hoofs or dewclaws; in the hippopotamus alone they are fully developed and the animal has a four-toed foot.
This sole, Smith states, softens in the spring of the year, when the snow is leaving the ground, a fresh layer of the integument taking its place. The rubber-like balls with which the dewclaws are provided are by no means useless; they project back below the horny part of the hoof, and Mr.
Behind this crescentic horn is a shallow concavity which gives the horny hoof a chance to get its hold. Both the main digits and the dewclaws terminate in black, rubber-like, rounded and expanded soles, which are of great service in securing a firm footing on the shelving rocks and narrow ledges on which the animal travels with such ease.
Smith has actually observed the young captive goats supporting themselves solely on their dewclaws on the edge of a roof. There were certain trails leading over the hill slopes at Hui-yao which the gorals must have used continually, judging by the way in which these were worn.
In the Miocene the stature of the ancestors of the horse increased to that of a pony. The feet were still three-toed, but the side hoofs were now mere dewclaws and scarcely touched the ground. The evolution of the family was completed in the Pliocene. The middle toe was enlarged still more, the side toes were dropped, and the palm and foot bones which supported them were reduced to splints.
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