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Speaking generally, it may be said that human bones, where any were met with, occurred at all depths in the cave-mud and gravel, sometimes above and sometimes below those of the bear, elephant, rhinoceros, hyaena, etc. He preserved some few of them, however, which I have seen in the museum at Liege.

The bones, he suggested, may often have been rolled in the beds of such streams before they reached their underground destination. To the same agency the introduction of many land-shells dispersed through the cave-mud was ascribed, such as Helix nemoralis, H. lapicida, H. pomatia, and others of living species.

First, there was a period when the carbonate of lime was carried out gradually by springs; secondly, an era when engulfed rivers or occasional floods swept organic and inorganic debris into the subterranean hollows previously formed; and thirdly, there were such changes in the configuration of the region as caused the engulfed rivers to be turned into new channels, and springs to be dried up, after which the cave-mud, breccia, gravel, and fossil bones would bear the same kind of relation to the existing drainage of the country as the older valley-drifts with their extinct mammalian remains and works of art bear to the present rivers and alluvial plains.

In all the subterranean cavities, more than forty in number, explored by Schmerling, he only observed one cave, namely that of Chokier, where there were two regular layers of stalagmite, divided by fossiliferous cave-mud.