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A true crocodile, also, Crocodilus toliapicus, and another saurian more nearly allied to the gavial, accompany the above fossils; also the relics of several birds and quadrupeds. One of these last belongs to the new genus Hyracotherium of Owen, of the hog tribe, allied to Chaeropotamus, another is a Lophiodon; a third a pachyderm called Coryphodon eocaenus by Owen, larger than any existing tapir.
With these are several species of the genus Oreodon, Leidy, uniting the characters of pachyderms and ruminants also; Eucrotaphus, another new genus of the same mixed character; two species of rhinoceros of the sub-genus Acerotherium, a Lower Miocene form of Europe before mentioned; two species of Archaeotherium, a pachyderm allied to Chaeropotamus and Hyracotherium; also Paebrotherium, an extinct ruminant allied to Dorcatherium, Kaup; also Agriochoerus, of Leidy, a ruminant allied to Merycopotamus of Falconer and Cautley; and, lastly, a large carnivorous animal of the genus Machairodus, the most ancient example of which in Europe occurs in the Lower Miocene strata of Auvergne, but of which some species are found in Pliocene deposits.
Where, then, must we look for its five-toed ancestor? If we follow down other lines of recent and tertiary Ungulata, the same question presents itself. The Pigs are traceable back through the Miocene epoch to the Upper Eocene, where they appear in the two well-marked forms of Hyopotamus and Chaeropotamus; but Hyopotamus appears to have had only two toes.
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