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Under this tissue is found with the Nos. 7, 8, and 9, the endosperm or perisperm, containing the gluten and the starch; soluble and insoluble albuminoids, that is to say, the flour.

In the monocotyledonous plants there is no milky juice; but the perisperm of the palms, which yields such sweet and agreeable milky emulsions, contains, no doubt, caseum.

Endospermic food-reserve has evident advantages over perispermic, and the latter is comparatively rarely found and only in non-progressive series. Seeds in which endosperm or perisperm or both exist are commonly called albuminous or endospermic, those in which neither is found are termed exalbuminous or exendospermic.

Those by which the sulphate of iron is precipitated of a green colour, like the real cinchona, the bark of the white willow, and the horned perisperm of the coffee-tree, do not on this account denote identity of chemical composition;* and that identity might even exist, without our concluding that the medical virtues were analogous.

In some plants the nucellus is not thus absorbed, but itself becomes a seat of deposit of reserve-food constituting the perisperm which may coexist with endosperm, as in the water-lily order, or may alone form a food-reserve for the embryo, as in Canna.