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Updated: October 18, 2026
As to the mere action of surrounding conditions, the same Professor remarks: "Any modification affecting the density of the soil might so far relate to the changes of limb-structure, as that a foot with a pair of small hoofs dangling by the sides of the large one, like those behind the cloven hoof of the ox, would cause the foot of Hipparion, e.g., and a fortiori the broader based three-hoofed foot of the Palæothere, to sink less deeply into swampy soil, and be more easily withdrawn than the more concentratively simplified and specialized foot of the horse.