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There are facts, however, which prove that all species are not sterile inter se, and that their characters when they are hybridised do not always segregate in Mendelian fashion. Exper. In the former cross he states that except for one or two characters there seemed to be no more tendency to variation in the F2 generation than in the F1. The statement is, however, open to criticism.
Motion is communicated from the way-shafts, w and w1, by the eccentrics, and the eccentric rods, e1 e2 e3 e4, and the levers and rods belonging thereto, to the short steam valve rocking shafts levers, f1 f2 f3 f4, and the exhaust valve rocking shafts, k1 k2 k3 k4, the bearings of which are carried on brackets above the valve chests, which, being furnished with tappet levers, raise and lower the valves.
In the F1, when the horned character in the female is only inherited from one side, the hereditary tendency is not enough to overcome the influence of the absence of the testis hormone and presence of the ovarian hormone, and so the horns do not develop.
In the F1 generation the males were horned, the females hornless. Here, with regard to the horned character, both sexes were of the same genetic composition, i.e. heterozygous, or if we represent the possession of horns by H, and their absence by h, both sexes were Hh.
The matrices falling from the magazine descend through the front channels and are received on the inclined belt F, on which they are carried over and guided on the upper rounding surface of the assembler entrance-block f1, by which they are guided downward in front of the star-wheel f2, which pushes them forward one after another.
Since the heterozygote in F1 was deeply pigmented, it is certain that a bird with only a small amount of pigment in its skin was a recessive resulting from incomplete segregation of the pigmented character. The pigment occurred chiefly in the skin of the abdomen and round the eyes, and also in the peritoneum and in the connective tissue of the abdominal wall.
Both the pigmented skin of the Silky and the colour in the plumage of the bankiva are dominant, so that all the offspring in F1 or the first generation are coloured fowls with pigmented skins. But in later generations I found that with regard to skin pigment there were no pure recessives.
Of the males one was a variate specially marked, about half-way between the F1 type and the Mallard parent. This, according to Phillips, was a segregate. The rest showed a range of variation but no distinct segregation.
The question is, what were the unit characters in the parent species? If the unit characters were very small and numerous, an individual in which all the characters of the Pintail existed together among the offspring of the hybrid mated with pure Pintail would be rare in proportion to the individuals presenting other combinations. The females were all alike and similar to F1 females.
Still more striking was the incomplete segregation in the plumage colour. The white of the Silky was recessive, all the birds of the F1 generation being fully coloured. In the F2 generation there were two recessive white cocks which when mature showed slight yellow colour across the loins.
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