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Even in this modified chordaea-theory the six fundamental organs of the chordula would retain their great value. The medullary tube would be originally a chemical sensory organ, a dorsal olfactory tube, taking in respiratory-water and food by the neuroporus in front and conveying them by the neurenteric canal into the primitive gut.

We will now compare the embryonic development of the two animals, and find to our great astonishment that the same embryonic form develops from the ovum of the Amphioxus as from that of the Ascidia a typical chordula.

As both the Tunicates and the Vertebrates develop from the same chordula, we may infer that there was a corresponding common ancestor of both stems. We may call this the Chordaea, and the corresponding stem-group the Prochordonia or Prochordata. We shall see presently how this conclusion is justified in the present condition of morphological science.

Not less important are the later embryonic forms of the coelomula, the chordula, etc. Thus the embryology of the Amphioxus and the Ascidia has so much increased our knowledge of man's stem-history that, although our empirical information is still very incomplete, there is now no defect of any great consequence in it.

We have first to answer the difficult and much-discussed question of the development of the Chordaea from the Gastraea; in other words, "How and by what transformations were the characteristic animals, resembling the embryonic chordula, which we regard as the common stem-forms of all the Chordonia, both Tunicates and Vertebrates, evolved from the simplest two-layered Metazoa?"