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In all cases the acquirement of sensitiveness by the mid-ribs of the leaves appears to stand in some close relation with the abortion of their laminae or blades. The latter, therefore, are intermediate in nature between twiners and tendril-bearers, and ought to be related to both.

A greater number of twiners revolve in a course opposed to that of the sun, or to the hands of a watch, than in the reversed course, and, consequently, the majority, as is well known, ascend their supports from left to right.

On the other hand, revolving internodes, without any other aid, suffice to give the power of climbing; so that it seems probable that leaf-climbers were in most cases at first twiners, and subsequently became capable of grasping a support; and this, as we shall presently see, is a great additional advantage.

We passed very many sites of old villages, which are easily known by the tree euphorbia planted round an umbelliferous one, and the sacred fig. One species here throws out strong buttresses in the manner of some mangroves instead of sending down twiners which take root, as is usually the ease with the tropical fig.

In those plants which still twine, but at the same time possess tendrils or sensitive petioles, as some species of Bignonia, Clematis, and Tropaeolum, it can readily be observed how incomparably better they grasp an upright stick than do simple twiners.

Nine species of Bignonia, selected by hazard, are here described, in order to show what diversity of structure and action there may be within the same genus, and to show what remarkable powers some tendrils possess. The species, taken together, afford connecting links between twiners, leaf-climbers, tendril-bearers, and root-climbers.

This is the case: thus the several leaf-climbing species of the Antirrhineae, of Solanum, Cocculus, and Gloriosa, have within the same family and even within the same genus, relatives which are twiners. In the genus Mikania, there are leaf-climbing and twining species. The leaf-climbing species of Clematis are very closely allied to the tendril-bearing Naravelia.

I believe it is not a fact that the ivy sucks the juice out of the trees it climbs upon, though it may much interfere with their growth. Its aerial rootlets are for support alone, as is the case with all climbers that are not twiners.

Most twining plants are adapted to ascend supports of moderate though of different thicknesses. In the open air, however, the Phaseolus twined round a support of the above thickness, but failed in twining round one 9 inches in diameter. Nevertheless, some twiners of the warmer temperate regions can manage this latter degree of thickness; for I hear from Dr.

The two great classes of twiners and of plants with sensitive organs, namely, leaf-climbers and tendril-bearers taken together, far exceed in number and in the perfection of their mechanism the climbers of the two first classes.