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Some species stridulate very loudly, so that when Mr. F. Smith caught a Trox sabulosus, a gamekeeper, who stood by, thought he had caught a mouse; but I failed to discover the proper organs in this beetle. I am indebted to Mr. In Oryctes it is seated on the pro-pygidium; and, according to the same entomologist, in some other Dynastini, on the under surface of the elytra.
The largest are the three thoracic ganglia and the eleventh. After ascertaining these facts, I remembered Swammerdam's investigations into the grub of the Monoceros, our Oryctes nasicornis. I consulted the venerable volume.
The black Cricket, with his red-braided thighs, is the infallible label of the Yellow-winged Sphex; the larva of Oryctes nasicornis tells us of the Garden Scolia as certainly as the best description; the Cetonia-grub proclaims the Two-banded Scolia and the larva of the Anoxia announces the Interrupted Scolia.
I select from my heap of garden-mould, that inexhaustible mine, two larvae of the Rhinoceros Beetle, Oryctes nasicornis, about one-third full-grown, so that their size may not be out of proportion to the Scolia's. It is in fact almost identical with the size of the Cetonia. I paralyse one of them by giving an injection of ammonia in the nerve-centres.
Nevertheless, let us put the precursor of the Scoliae to the test. What did she do? Being capable of everything, she did a bit of everything. Among its descendants were innovators who developed a taste for tunnelling in sand and vegetable mould. There they encountered the larvae of the Cetonia, the Oryctes, the Anoxia, succulent morsels on which to rear their families.
This precursor, modified by varying circumstances, is supposed to have subdivided herself into ramifications, one of which, digging into vegetable mould and preferring the Cetonia to any other game inhabiting the same heap, became the Two-banded Scolia; another, also addicted to exploring the soil, but selecting the Oryctes, left as its descendant the Garden Scolia; and a third, establishing itself in sandy ground, where it found the Anoxia, was the ancestress of the Interrupted Scolia.
Apart from a few details, which are of no importance here, we may repeat this sketch of the Anoxia-grub and we shall have, if the size be at least quadrupled, a picture of the larva of Oryctes nasicornis, the monstrous prey of the Garden Scolia. Its general appearance is the same: there is the same exaggeration of the belly; the same hook-like curve; the same incapacity for standing on its legs.
The Garden Scolia attacks Oryctes nasicornis; the Two-banded Scolia the Cetonia; the Interrupted Scolia the Anoxia. All three operate below ground, under the most unfavourable conditions; and all three have for their victim a larva of one of the Scarabaeidae, which, thanks to the exceptional arrangement of its nerve-centres, lends itself, alone of all larvae, to the Wasp's successful enterprises.
It includes the largest of our Beetles, the common Rhinoceros Beetle, or Oryctes nasicornis. I find some who have been recently liberated, whose wing-cases, of a glossy brown, now see the sunlight for the first time; I find others enclosed in their earthen shell, almost as big as a Turkey's egg. More frequent is her powerful larva, with its heavy paunch, bent into a hook.
I note the presence of a second bearer of the nasal horn, Oryctes Silenus, who is much smaller than her kinswoman, and of Pentodon punctatus, a Scarabaeid who ravages my lettuces. But the predominant population consists of Cetoniae, or Rosechafers, most of them enclosed in their egg-shaped shells, with earthen walls encrusted with dung.
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