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Thus there is a vast negative difference between the Cainozoic and the Mesozoic mammalian faunae of Europe. But there is a still more important positive difference, inasmuch as all these Mammalia appear to be Marsupials belonging to Australian groups, and thus appertaining to a different distributional province from the Eocene and Miocene marsupials, which are Austro-Columbian.
And the imperfect knowledge we have of the ancient mammalian population of our earth leads to the belief that certain of its types, such as that of the 'Marsupialia', have persisted with correspondingly little change through a similar range of time.
Uterine and mammary functions are generally regarded as essentially female characteristics, and are included in the popular idea of the sex of woman. Scientifically, of course, they are not at all necessary or universal features of the female sex, but are peculiar to the mammalian class of Vertebrates in which they have been evolved.
Like the reptiles they lay tough-coated eggs and have a single outlet for the excreta, and they have a reptilian arrangement of the bones of the shoulder-girdle; like the mammals, they have a coat of hair and a four-chambered heart, and they suckle the young. Even in their mammalian features they are, as the careful research of Australian zoologists has shown, of a transitional type.
The fact that this immense fauna of Miocene Arctogaea is now fully and richly represented only in India and in South Africa, while it is shrunk and depauperised in North Asia, Europe, and North America, becomes at once intelligible, if we suppose that India and South Africa had but a scanty mammalian population before the Miocene immigration, while the conditions were highly favourable to the new comers.
His favorite thesis included the origin of mammalian life and of man himself in southernmost South America, with, as incidents, the belief that the mammalian-bearing strata of South America were of much greater age than the strata with corresponding remains elsewhere; that in South America various species and genera of men existed in tertiary times, some of them at least as advanced as fairly well advanced modern savages; that there existed various land bridges between South America and other southern continents, including Africa; and that the ancestral types of modern mammals and of man himself wandered across one of these bridges to the old world, and that thence their remote descendants, after ages of time, returned to the new.
In place of the petty phalangers and pouched ant-eaters of the oolitic period, our tertiary strata in the larger continents show us a rapid and extraordinary development of the mammalian race into monstrous creatures, some of them now quite extinct, and some still holding their own undisturbed in India, Africa, and the American prairies.
Why then should not the reduction of equally useless, more wasteful, and perhaps positively dangerous wings be also due to an economy which has become advantageous to bird and reptile alike through the absence of the mammalian rivals whose places they are evidently being modified to fill?
One fancies every living thing as not only returning its mineral elements to the soil, but as in some subtle way leaving its vital forces also, and thus contributing to the impalpable, invisible store-house of vital energy of the globe. At first among the mammalian tribes there was much muscle and little brains.
To cite but a single illustration, the marsupial order, which is the dominant mammalian type of the living fauna of Australia to-day, existed in Europe and died out there in the tertiary age. Hence a future geologist might think the Australia of to-day contemporaneous with a period in Europe which in reality antedated it by perhaps millions of years.
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