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When the crimson threads appear in the scaly buds the staminate catkins are lengthening, and soon the high wind shakes the golden pollen over all the copse. These flowers which appear before the leaves all depend upon the wind for their fertilization. That is why they come before the leaves. And there is always wind enough to meet all their needs. March is a masculine month.
In the case of the hepatica acutiloba, however, it has been found that staminate flowers grow on one plant and pistillate flowers on another, hence insects are essential to the perpetuation of this species. After bringing us the trilliums and hepaticas in numbers, Nature pauses.
But if one set out a bed of Champions another fine variety well apart from any staminate kind, it would blossom profusely, but produce no fruit. When I was a boy, Hovey's Seedling was the great strawberry of the day, and marvellous stories were told of the productiveness of the plants and the size of the berries.
All danger of barrenness in pistillate kinds is removed absolutely by planting staminate sorts in the same bed. In nurserymen's catalogues pistillate varieties are marked "P.," and the purchaser has merely to set out the plants within a few feet of some perfect flowering kind to secure abundant fruit. As a result of much experience, I will now make some suggestions as to varieties.
How well I remember the disappointment and wrath of people who bought the plants at a high price, and set them out with no staminate varieties near to fertilize the pistillate blossoms. Expectations were raised to the highest pitch by profuse blossoming in May, but not a berry could be found the ensuing June.
Then they were wee golden-green buds of pistillate flowers, set high on the uppermost branch tips that the pollen from the tree's own staminate blooms might miss them in its flight down the wind and thus avoid in breeding. If they miss fertilization altogether they fall off.
Meanwhile the staminate catkins of the hazel are lengthening and the pistillate buds are swelling, as the sun presses farther northward at the dawn and the dusk of each day, pushing back the gray walls of the cañon of night, that the river of day may flow full and free. This year some of the aspens heralded the spring.
In some flowers this separation is effected, as I have shown, by their maturing at different periods; in others, as in the iris, by mere mechanical means; while in a long list of plants, as in the willow, poplar, hemp, oak, and nettle, the cross-fertilization is absolutely necessitated by the fact of the staminate and stigmatic flowers being either separated on the same stalk or on different plants, the pollen being carried by insects or the wind.
Many of these cases return regularly in each generation, or may at least be seen from time to time in the same strains. Proliferation of the inflorescence is very common and changes in the position of staminate and pistillate flowers are not rare. We find starting points for new investigations in almost any teratological structure.
Youth will have its way, and in this case it seems to seize on the first sap that gets as far as the topmost branch tips, compelling it to the nourishing of the young cones before it can go to the making of new leaves or even of the crop of staminate and pistillate blossoms for the ensuing summer.
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