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The Insemineae include forms in which the nucellus is not developed, and therefore there can be no seed. The plants included are, however, mainly well-established parasites, and the absence of nucellus is only one of those characters of reduction to which parasites are liable.

In some plants the nucellus is not thus absorbed, but itself becomes a seat of deposit of reserve-food constituting the perisperm which may coexist with endosperm, as in the water-lily order, or may alone form a food-reserve for the embryo, as in Canna.

The development of the ovule, which represents the macrosporangium, is very similar to the process in Gymnosperms; when mature it consists of one or two coats surrounding the central nucellus, except at the apex where an opening, the micropyle, is left. The nucellus is a cellular tissue enveloping one large cell, the embryo-sac or macrospore.

Ovular characters determine the grouping in the Dicotyledons, van Tieghem supporting the view that the integument, the outer if there be two, is the lamina of a leaf of which the funicle is the petiole, whilst the nucellus is an outgrowth of this leaf, and the inner integument, if present, an indusium.

In some cases the embryo or the embryo-sac sends out suckers into the nucellus and ovular integument.

Isolated cases show that any of the cells within the embryo-sac may exceptionally form an embryo, e.g. the synergidae in species of Mimosa, Iris and Allium, and in the last-mentioned the antipodal cells also. In a species of Allium, embryos have been found developing in the same individual from the egg-cell, synergids, antipodal cells and cells of the nucellus.