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In 1859, there appeared to be a very sharp and clear hiatus between vertebrated and invertebrated animals, not only in their structure, but, what was more important, in their development.

A general review of the whole field of Biology not only of the Natural History of the present but of the geological record in relation to the known laws and the various main theories of the evolutionary process will be taken, and in addition some special department, either the Comparative Anatomy of the Vertebrata chiefly, or of the plants chiefly, or of several Invertebrated groups chiefly, will be exhaustively worked out in relation to these speculations.

The discrepancy between the two conceptions is surely overpowering, when we allow ourselves to see the whole matter in a steady and rational light. There is, also, the striking fact of an ascertained historical progress of plants and animals in the order of their organization; marine and cellular plants and invertebrated animals first, afterwards higher examples of both.

But, although palaeontology is a comparatively youthful scientific speciality, the mass of materials with which it has to deal is already prodigious. In the last fifty years the number of known fossil remains of invertebrated animals has been trebled or quadrupled.