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Professor Huxley called attention to this subject in a brief address to the London Ethnological Society in 1869. After stating the case, he presented the following queries and suggestions: “The Austro-Columbian fauna, as a whole, therefore, existed antecedently to the glacial epoch. Did man form part of that fauna?
It is unfortunate that we have no knowledge of the Miocene mammalian fauna of the Australian and Austro-Columbian provinces; but, seeing that not a trace of a Platyrrhine Ape, of a Procyonine Carnivore, of a characteristically South-American Rodent, of a Sloth, an Armadillo, or an Ant-eater has yet been found in Miocene deposits of Arctogaea, I cannot doubt that they already existed in the Miocene Austro-Columbian province.
It is unfortunate that we have no knowledge of the Miocene mammalian fauna of the Australian and Austro-Columbian provinces; but, seeing that not a trace of a Platyrrhine Ape, of a Procyonine Carnivore, of a characteristically South-American Rodent, of a Sloth, an Armadillo, or an Ant-eater has yet been found in Miocene deposits of Arctogaea, I cannot doubt that they already existed in the Miocene Austro-Columbian province.
Assuming this supposition to be correct, we should have to look in the human population of America, as in the fauna generally, for an indigenous or Austro-Columbian element, and an immigrant or ‘Arctogeal’ element.” He then suggests that the Esquimaux may now represent the immigrant element, and the old Mexican and South American race that which was indigenous, and that the “Red Indians of North America” may have appeared originally as a mixture of these two races.
Thus there is a vast negative difference between the Cainozoic and the Mesozoic mammalian faunae of Europe. But there is a still more important positive difference, inasmuch as all these Mammalia appear to be Marsupials belonging to Australian groups, and thus appertaining to a different distributional province from the Eocene and Miocene marsupials, which are Austro-Columbian.
But Austro-Columbia presents difficulties from which Australia is free; Camelidae and Tapiridae are now indigenous in South America as they are in Arctogaea; and, among the Pliocene Austro-Columbian mammals, the Austro-Columbian genera Equus, Mastodon, and Machairodus are numbered. Are these Postmiocene immigrants, or Praemiocene natives? Cunningham sent over to me some time ago from Patagonia.
But Austro-Columbia presents difficulties from which Australia is free; Cantelidoe and Tapirdoe are now indigenous in South America as they are in Arctogaea; and, among the Pliocene Austro-Columbian mammals, the Arctogaeal genera Equus, Mastodon, and Machairodus are numbered. Are these Postmiocene immigrants, or Praemiocene natives? Cunningham sent over to me some time ago from Patagonia.
Thus there is a vast negative difference between the Cainozoic and the Mesozoic mammalian faunae of Europe. But there is a still more important positive difference, inasmuch as all these Mammalia appear to be Marsupials belonging to Australian groups, and thus appertaining to a different distributional province from the Eocene and Miocene marsupials, which are Austro-Columbian.
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