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On comparing a thin transverse slice of such a petiole with one from an older leaf growing close beneath, which had not clasped anything, its diameter was found to be fully doubled, and its structure greatly changed. In two other petioles similarly compared, and here represented, the increase in diameter was not quite so great.

There is no such movement in any leaf- climber, with the exception of an occasional trace of it in the petioles of Tropaeolum tricolorum. On the other hand, the tendrils of all tendril-bearing plants, contract spirally after they have caught an object with the following exceptions. Firstly, Corydalis claviculata, but then this plant might be called a leaf-climber.

In all cases, if the sticks are taken away, the petioles continue to move during many hours afterwards; so they do after a slight rubbing; but they become straight again, after about a day's interval, that is if the flexure has not been very great or long continued. The graduated difference in the extension of the sensitiveness in the petioles of the above-described species deserves notice.

Leaves alternate, with very short petioles, at the base of which is a pair of short spines, and a small tuft of wool in the axil; blade 3 in. long by 2 in. broad, soft, fleshy, shining green. Flowers semi-transparent, white, in terminal panicles; sepals and petals ¾ in. long by ¼ in. wide; stamens in a large, spreading cluster, white, with yellow anthers.

The young petioles, which alone are sensitive, have their ends bowed a little downwards, so as to be in a slight degree hooked; ultimately the whole leaf, if it catches nothing, becomes level.

The leaves are so small compared with those of most of the other species, that the petioles at first seem ill-adapted for clasping. Nevertheless, the main service of the revolving movement is to bring them into contact with surrounding objects, which are slowly but securely seized.

That this saving of the stem is really an advantage to climbing plants, I infer from the species that still twine but are aided by clasping petioles or tendrils, generally making more open spires than those made by simple twiners. Moreover, the plants thus aided, after taking one or two turns in one direction, generally ascend for a space straight, and then reverse the direction of their spire.

Hence, perhaps, it is that the petioles have acquired a high degree of sensitiveness as a compensation for the little power of movement in the shoots. The petioles are bowed downwards, and have the same general hook-like form as in C. viticella. The medial petiole and the lateral sub- petioles are sensitive, especially the much bent terminal portion.

On February 5 all the petioles which had been drawn into the burrows beneath a Robinia, were pulled up; and after an interval of eleven days, 35 petioles had been again dragged in, 19 by the base, and 16 by the apex. Taking these two lots together, 56 per cent. were drawn in by the base, and 44 per cent. by the apex.

They act in nearly the same manner as would a man, who had to close a cylindrical tube with different kinds of leaves, petioles, triangles of paper, etc., for they commonly seize such objects by their pointed ends. But with thin objects a certain number are drawn in by their broader ends.