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The tree-creepers do not possess melodious, or at any rate mellow notes, although in so numerous a family there is great variety of tone, ranging from a small reedy voice like the faint stridulation of a grasshopper, to the resounding, laughter-like, screaming concerts of Homorus, which may be heard distinctly two miles away.

The allied Homorus lophotis builds a nest equally large, but with a small cavity for the eggs inside, and outwardly resembling a gigantic powder-flask, lying horizontally among the lower branches of a spreading tree.

Some have a spiral passage-way leading from the entrance to the nest cavity, and the cavity is in many cases only large enough to accommodate the bird; but in the gigantic structure of Homorus gutturalis it is so large that, if the upper half of the nest or dome were removed, a condor could comfortably hatch her eggs and rear her young in it. This nest is spherical.

Coryphistera resembles the lark and pipit in its habits; Cinclodes, the wagtail; Geobates a Saxicola; Limnornis lives in reed beds growing in the water; Henicornis in reed beds growing out of the water; and many other ground species exist concealed in the grass on dry plains; Homorus seeks its food by digging in the loose soil and dead leaves about the roots of trees; while Geo-sitta, Furnarius, and Upercerthia obtain a livelihood chiefly by probing in the soil.

The habit varies somewhat in the cachalote, a Patagonian species of the genus Homorus, about the size of the missel-thrush. Old and young birds live in a family together, and at intervals, on any fine day, they engage in a grand screaming contest, which may be heard distinctly at a distance of a mile and a half.