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In the polypetalous forms progression from hypogyny to epigyny is generally recognized, and where dorsiventrality with insect-pollination has been established, a dominant group has been developed as in the Leguminosae. The starting-point of the class, however, and the position within it of apetalous families with frequently unisexual flowers, have provoked much discussion.

The forest trees on the ironstone ridges were stringy-bark, and on the grassy hills box, Moreton Bay ash, and a tree belonging to the natural order Leguminosae, with axillary racemes of white apetalous flowers, long, broad, flat, many-seeded legumes, large, bipinnate leaves, leaflets oval, one inch long, and having dark fissured bark; on the flat stiff soil grew ironbark, apple-tree, and another species of Angophora, with long lanceolate leaves, seed vessels as large as the egg of a common fowl and a smooth yellow bark.

Apetalous flowers may be considered as another form of monstrosity, and in Salpiglossis sinuata such a variety without a corolla made its appearance in the year 1892 in the nursery of Vilmorin. It appeared suddenly, yielded a good crop of seed and was constant from the outset, without any sign of vicinism or impurity.